Osteoglossiformes (Bony tongues)
Elopiformes (Tarpon / Tenpounders)
- Elops sp.
- Esocelops cavifrons
- Protarpon (Megalops) oblongus
- Protarpon (Megalops) priscus
- Promegalops sheppeyensis
- Promegalops signeuxae
Aulopiformes (Grinners & Lizardfish)
Beryciformes (Soldier/Pricklefish)
Scombriformes (Mackerel & Tuna)
- Eocoelopoma colei
- Eocoelopoma curvatum
- Eocoelopoma gigas
- Micrornatus hopwoodi
- Scombramphodon crassidens
- Scombramphodon sheppeyensis
- Duplexdens (Scombrinus) macropomus
- Scombrinus nuchalis
- Sphyraenodus priscus
- Tamesichthys decipiens
- Wetherellus brevior
- Wetherellus cristatus
- Wetherellus longior
- Woodwardella patellifrons
- Palaeocybium proosti
- Bramoides brieni
- Goniocranion arambourgi
- Undescribed Scombrid #1
- Undescribed Scombrid #2
Carangiformes (Jack Mackerels)
- Enniskillenus radiatus
- Cylindracanthus rectus
- Hemirhabdorhynchus elliotti
- Acestrus elongatus
- Acestrus ornatus
- Aglyptorhynchus sulcatus
- Aglyptorhynchus venablesi
- Xiphiorhynchus priscus
Pycnodus toliapicus
Scientific Classification
Class: Actinopterygii (Ray-finned Fish)
Order: Pycnodontiformes (Pycnodonts)
Family: Pycnodontidae
Genus: Pycnodus [Pyke-node-us]
Species: toliapicus (Agassiz 1839)
There are a number of specimens known of this species of Pycnodus which a number represented in the collection of the Natural History Museum, London – most notably a matching pair of lower jaw splenials.
- Multiple rows to crushing teeth which are rounded and elongated (1)
- Each tooth features a flattened surface with gently rounded edges (2)
- Marginal teeth reduce in size and become more rounded in shape with some pitting and texture. The second column of teeth measures about 50% of the size of the largest teeth which differs from P.bowerbanki which remains more elongated and about 75% of the length of the largest teeth (3)


3D scans undertaken by Tom Sermon with permission. From the collections of the Natural History Museum, London

Casier, E. (1966) Fauna ichthyologique du London Clay. British Museum Natural History London, 1966, 1-496
Monsch, K (2005) Revision of the scombroid fishes from the Cenozoic of England
Raynor, Mitchell, Clouter (2009) London Clay Fossils of Kent and Essex